The principle of faunal succession, also known as the law of faunal succession, is based on the observation that sedimentary rock strata contain

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

ized

flora Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' ...

and

fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is ''funga''. Flora, fauna, funga and other forms of life are collectively referred to as ''Biota (ecology ...

, and that these fossils succeed each other vertically in a specific, reliable order that can be identified over wide horizontal distances. A fossilized

Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While the ...

bone will never be found in the same stratum as a fossilized

Megalosaurus ''Megalosaurus'' (meaning "great lizard", from Greek , ', meaning 'big', 'tall' or 'great' and , ', meaning 'lizard') is an extinct genus of large carnivorous theropod dinosaurs of the Middle Jurassic period (Bathonian stage, 166 million years ...

, for example, because neanderthals and megalosaurs lived during different

geological periods The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochron ...

, separated by many millions of years. This allows for strata to be identified and dated by the fossils found within. This principle, which received its name from the English

geologist A geologist is a scientist who studies the solid, liquid, and gaseous matter that constitutes Earth and other terrestrial planets, as well as the processes that shape them. Geologists usually study geology, earth science, or geophysics, althoug ...

William Smith, is of great importance in determining the relative age of rocks and strata. The fossil content of rocks together with the

law of superposition The law of superposition is an axiom that forms one of the bases of the sciences of geology, archaeology, and other fields pertaining to geological stratigraphy. In its plainest form, it states that in undeformed stratigraphic sequences, the ...

helps to determine the time sequence in which sedimentary rocks were laid down.

Evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...

explains the observed faunal and floral succession preserved in rocks. Faunal succession was documented by Smith in England during the first decade of the 19th century, and concurrently in France by Cuvier (with the assistance of the mineralogist Alexandre Brongniart). Archaic biological features and organisms are succeeded in the

fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

by more modern versions. For instance,

paleontologists Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of foss ...

investigating the evolution of

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...

s predicted that feathers would first be seen in primitive forms on flightless predecessor organisms such as

feathered dinosaur A feathered dinosaur is any species of dinosaur possessing feathers. While this includes all species of birds, there is a hypothesis that many, if not all non-avian dinosaur species also possessed feathers in some shape or form. It has been su ...

s. This is precisely what has been discovered in the fossil record: simple feathers, incapable of supporting flight, are succeeded by increasingly large and complex feathers. In practice, the most useful diagnostic species are those with the fastest rate of species turnover and the widest distribution; their study is termed

biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of B ...

, the science of dating rocks by using the

s contained within them. In Cenozoic strata, fossilized tests of

foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly ...

are often used to determine faunal succession on a refined scale, each biostratigraphic unit (

biozone In biostratigraphy, biostratigraphic units or biozones are intervals of geological strata that are defined on the basis of their characteristic fossil taxa, as opposed to a lithostratigraphic unit which is defined by the lithological properties ...

) being a geological stratum that is defined on the basis of its characteristic fossil

taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

. An outline microfaunal zonal scheme based on both foraminifera and

ostracod Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typi ...

a was compiled by M. B. Hart (1972). Earlier fossil life forms are simpler than more recent forms, and more recent fossil forms are more similar to living forms (principle of faunal succession).''Evolutionary Analysis'', 4th Edition. p 61.

References

{{Geologic Principles Paleontological concepts and hypotheses Dating methods Stratigraphy de:Biostratigraphie#Das Prinzip der Fossilfolge

See also

References